The story takes place before Puss met Shrek, when he was a swashbuckling hero who protected the innocent. A bunch of old pub thugs tell him that two murderous outlaws called Jack and Jill have discovered an ancient power that can destroy the world. Puss then sets off on a journey with his old friend Humpty Dumpty who also introduces him to Kitty Softpaws, a sly black cat who takes an interest in Puss's journey. With his new sidekicks, Puss sets off on his most adventurous and dangerous journey ever.
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